The eldest son of the eldest branch was born noble and had a lot of responsibilities. He was nurtured as the heir since he was young.
With eugenics and education in mind, they chose the eldest daughter-in-law carefully to ensure that the descendants of the eldest branch would not have IQ problems.
In Mu Ci's generation, due to physical reasons and the fact that he had long been judged to not live long, the responsibility of the eldest branch fell on Mu En.