Never in her wildest dreams did Audrey think that she would be embraced by the person that she was ordered to kill several years ago.
Together with Lira and Ephemera, they went to hunt the Half-Elf under the Pope's orders. Unfortunately, this order backfired on them and Lira and Ephemera were caught after facing off against William.
They were not aware that when his two sisters returned, their wombs had already been tainted with William's seed, bearing his mark.
After Lira and Ephemera, Melody was the next one to fall. Her sister, who was one of the kindest people she had met in her life, fell in love with the Half-Elf, and was soon conquered by him as well.
As for Shana, the Virtuous Maiden of Prudence became William's collaborator and looked after her sisters, who had already sided with him.
After thinking back, she realized that it was only her and Cherry that sided with the Pope until the latter betrayed their trust.