Lindo was both happy and flustered as he saw his father entering through the gates.
Happy because he didn't get to see his dad very often due to his job, and flustered because his dad disapproved of his relationship with Manitola.
It was mostly because his father wanted him to marry a female lion instead of someone from a different tribe.
It wasn't that he was against such type of practices but he wanted healthy descendants, he wanted to be a grandfather and it was unknown if the hybrid between a fox and a lion could even survive.
Of course, as the people on the continents evolved and gained a human form interbreeding was possible but mostly besides tribes that were similar between them as the dog and wolf tribes.